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Magasanik B

Jia M, Li Z, Lou Y, et al

Interestingly, the expansion of Clade 2 GS enzymes, which preceded that of the GS-like effectors, is common to cyst nematodes, root-knot nematodes, and indeed all endoparasitic nematode species within the Tylenchida (including those that do not establish a feeding site within the host)